Which birds use their beaks upside down?

toucans
flamingos
storks
penguins
Flamingos filter-feed on brine shrimp and blue-green algae. Their beaks are specially adapted to separate mud and silt from the food they eat, and are uniquely used upside-down. The filtering of food items is assisted by hairy structures called lamellae which line the mandibles, and the large rough-surfaced tongue.
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toucans
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3%
26
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71%
7%
0%
14
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63%
12%
1%
66
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17
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15
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193
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9%
11
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19%
6%
324
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40%
4%
5%
14935
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39%
12%
4%
481
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37%
25%
0%
16
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33%
0%
14%
21
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6%
419
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24%
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25
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27%
12%
30%
33
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6%
4%
63
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26%
17%
8%
23
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25%
8%
16%
12
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72%
14%
10%
2%
50