Which birds use their beaks upside down?

toucans
flamingos
storks
penguins
Flamingos filter-feed on brine shrimp and blue-green algae. Their beaks are specially adapted to separate mud and silt from the food they eat, and are uniquely used upside-down. The filtering of food items is assisted by hairy structures called lamellae which line the mandibles, and the large rough-surfaced tongue.
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